Medical School: University of California, San Francisco (1987-1992)
Residency: Internal Medicine, University of California, San Francisco (1992-1995)
Fellowship: Gastroenterology, Stanford University,Palo Alto, California (1995-2002)

1989 – 1992 Clerkship honors in Biochemistry Research, Infectious Diseases, Surgery, and Internal Medicine and Trauma Surgery subinternships – UCSF Medical School

1990 – 1991 Howard Hughes Medical Research Training Fellowship, laboratory of Dr. Peter Walter, Department of Biochemistry, UCSF

1991 – 1992 Howard Hughes Medical Institute Fellowship for continued medical educational support

1997 Howard Hughes Medical Institute Physician Postdoctoral Fellowship

2002 Commendation for service, Veterans Affairs Palo Alto Health Care System